Degrees

• B.S., Botany, Weber State College, 1979
• M.S., Biological Sciences, Northern Arizona University, 1981
• Ph.D., Genetics, Iowa State University, 1984

Research Interests

Research emphasis is on the genetic control of wheat end-use quality. Specific studies included the effects of alien genes on wheat quality, improving wheat quality by manipulating glutenin genes, and the development of reduced amylose and amylose-free wheats.

Major Project Activities

• Wheat molecular and biochemical genetics;
• Expression of alien genes in wheat;
• Development of amylose-free and reduced amylose wheats.

Selected Publications

• Baenziger, P. S. R. A. Graybosch, L. A. Nelson, T. Regassa,R. N. Klein, D. D. Baltensperger, D. Santra, A. M. H. Ibrahim, W. Berzonsky, J. M. Krall, L. Xu, S. N. Wegulo, M. Bernards, Y. Jin, J. Kolmer, J. H. Hatchett, Ming-shun Chen, and Guihua Bai. Registration of 'NH03614 CL' wheat. Journal of Plant Registrations. In press.
   
• Graybosch, R.A. and C. J. Peterson. 2010. Genetic improvement in winter wheat yields in the Great Plains of North America, 1959-2008. Crop Science. In press.
   
• Hansen, L.E., D.S. Jackson, R.L. Wehling, J.D. Wilson and R.A. Graybosch. 2010. Functionality of native tetraploid wheat starches: Effects of waxy loci alleles and amylose concentrations in blends. Journal of Cereal Science. 52: 39-45.
   
• Hansen, L.E., D.S. Jackson, R.L. Wehling and R.A. Graybosch. 2010. Functionality of chemically modified wild-type, partial waxy and waxy starches from tetraploid wheats. Journal of Cereal Science. 51:409-414.
   
• P. S. Baenziger, R. A. Graybosch, L. A. Nelson, R. N. Klein, D. D. Baltensperger, L. Xu, S. N. Wegulo, J. E. Watkins, Y. Jin, J. Kolmer, J. H. Hatchett, M.-S. Chen, and G. Bai. 2009. Registration of ‘Camelot’ wheat. Journal of Plant Registrations 3: 256-263.
   
• Graybosch, R.A. 2009. A review of the book: Wheat Chemistry and Technology. Journal of Cereal Science. 50: 304.
   
• Pierucci, V. R. M., M. Tilley, R. A. Graybosch, A. E. Blechl, S. R. Bean, and K. A. Tilley. Effects of Over-Expression of High Molecular Weight Glutenin Subunit 1Dy10 on Wheat Tortilla Properties. J. Agric. Food Chem. 57:6318–6326. 2009.
   
• Lan Guan, P. A. Seib, R. A. Graybosch, S. Bean, and Yong-Cheng Shi. 2009. Dough rheology and wet milling of hard waxy wheat flours. J. Agric. Food Chem. 57:7030-7038.
   
• Dowell, F.E., E.B. Maghirang, R.A. Graybosch, W.A. Berzonsky, and S.R. Delwiche. Selecting and sorting waxy wheat kernels using near-infrared spectroscopy. Cereal Chem. 86(3):251-255. 2009.
   
• Yongliang Liu, S. R. Delwiche and R. A. Graybosch. Two-dimensional correlation analysis of near infrared spectral intensity variations of ground wheat. Journal of Near Infrared Reflectance Spectroscopy 17: 41-50. 2009.
   
• Graybosch, R.A. and D.D. Baltensperger. Evaluation of the waxy endosperm trait in proso millet (Panicum mileaceum L.). Plant Breeding. 128: 70-73. 2009.
   
• Graybosch, R.A., C.J. Peterson, P. S. Baenziger, D. D. Baltensperger, L. A. Nelson, Y. Jin, J. Kolmer, B. Seaborn, R. French, G. Hein, T.J. Martin, B. Beecher, T. Schwarzacher and P. Heslop-Harrison. Registration of ‘Mace’ hard red winter wheat. Journal of Plant Registrations. 3: 51-56. 2009.
   
• Shubing Liu, Shibin Cai, R.A. Graybosch, Cuixia Chen and Guihua Bai. Quantitative trait loci for resistance to pre-harvest sprouting in U.S. hard white winter wheat Rio Blanco. Theoretical and Applied Genetics. 117:691–699. 2008.
   
• Saito, M., P. Vrinten, G. Ishikawa, R.A. Graybosch and T. Nakamura. A novel codominant marker for selection of the null Wx-B1 allele in wheat breeding programs. Molecular Breeding 23: 209-217. 2008.
   
• Baenziger, P.S., R.A. Graybosch, A. Ibrahim, D.D. Baltensperger, L.A. Nelson, Yue Jin, Ming Shun Chen, Guihua Bai, B.S. Beecher, J.H. Hatchett. Release of NE01643 hard red winter wheat. Journal of Plant Registrations. 2:36-42. 2008.
   
• Sarath, G., R. B. Mitchell, S. E. Sattler, D. Funnell, J. F. Pedersen, R. A. Graybosch and K. P. Vogel. Opportunities and roadblocks in utilizing forages and small grains for liquid fuels. Journal of Industrial Microbiology and Biotechnology. 35: 343-354. 2008.
   
• Heyduck, R.F., D. D. Baltensperger, L. A. Nelson, R. A. Graybosch. Assessing the stability of agronomic traits in waxy proso (Panicum miliaceum L.) in the central Great Plains. Crop Sci. 48:741-748. 2008.
   
• Pedersen, J.F., Graybosch, R.A., Funnell, D. Occurrence of the waxy alleles wxa and wxb in waxy sorghum plant introductions and their effect on grain thermal properties. Crop Science 47:1927-1933. 2007.
   
• Costa, J. M., C.A. Griffey, H. E. Bockelman, S. E. Cambron, A. Cooper, C. Gaines, R. A. Graybosch, A. Grybauskas, R. J. Kratochvil, Y. Jin, and D. Marshall. Registration of ‘Chesapeake’ Wheat. Journal of Plant Registrations; 1: 26 - 27. 2007.
   
• Graybosch, R.A. Potential for gene flow from cultivated wheat to weedy relatives in the Great Plains of North America. Proceedings of the OECD Conference on Biological Resources & Migration. In press. (conference proceedings).
   
• Graybosch, R.A. Grain Crops: Overview. Encyclopedia of Grain Science. In Press.
   
• Graybosch, R.A., N. Ames, P. S. Baenziger, and C. J. Peterson. Genotypic and Environmental Modification of Asian Noodle Quality of Hard Winter Wheats. Cereal Chemistry 81: 19-25. 2004.
   
• Wesley, I.J., B. G. Osborne, R. S. Anderssen, D. R. Delwiche, and R. A. Graybosch A chemomoetric localization approach to the nir measurement of apparent amylose content of ground wheat. Cereal Chemistry 80: 462-467. 2003.
   
• Guo, G. D. S. Jackson, R. A. Graybosch, and A. M. Parkhurst. Wheat tortilla quality: impact of amylose content adjustments using waxy wheat flour. Cereal Chemistry 80: 427-436. 2003.
   
• Guo, G., D. S. Jackson, R. A. Graybosch, and A. M. Parkhurst. Asian salted noodle quality: impact of amylose content adjustments using waxy wheat flour. Cereal Chemistry 80:437-445. 2003.
   
• Kim, W., Johnson, J.W., Graybosch, R.A. and Gaines, C.S. Physicochemical properties and end-use quality of wheat starch as a function of waxy protein alleles. Journal of Cereal Science. 37: 195-204. 2003.
   
• Graybosch, R.A., E. Souza, W. Berzonsky, P.S. Baenziger and O.K. Chung. Functional properties of waxy wheat flours: genotypic and environmental effects. J. Cereal Science 38: 69-76. 2003.
   
• Kumlay, A., P.S. Baenziger, K. Gill, D.R. Shelton, R.A. Graybosch, A. Lukaszewski and D. Wesenberg. Understanding the effect of rye chromatin in wheat. Crop Science 43: 1643-1651. 2003.
   
• Delwiche, S.R. and R.A. Graybosch. An examination of spectral pretreatments for PLS calibration for chemical and physical properties of wheat. Appl. Spectrosc. 57:1517-1527. 2003.
   
• Pirgozliev, V.R., S.P. Rose and R.A. Graybosch. Energy and amino acid availability to chickens of waxy wheat. Archiv fur Geflugelkunde. 66: 108-113. 2002.
   
• Souza, E., R.A. Graybosch and M. Guttiera. Breeding wheat for improved milling and baking quality. Journal of Crop Production 5: 39-74. 2002.
   
• Delwiche, S.R. and Graybosch, R.A. Identification of waxy wheat by near-infrared reflectance spectroscopy. Journal of Cereal Science 35: 29-38. 2002.
   
• Waniska, R.D., Graybosch, R.A. and Adams, J.L. Effect of Partial Waxy Wheat on Processing and Quality of Wheat Flour Tortillas. Cereal Chemistry 79: 210-214. 2002.
   
• Delwiche, S.R., Graybosch, R.A., Nelson, L. and Hruschka, W.R. Environmental effects on developing wheat as sensed by near-infrared reflectance of mature grains. Cereal Chemistry. 79:885-891. 2002.