Robert Graybosch
FYH 251
Lincoln NE 68583-0937



Robert Graybosch

Emeritus Professor

Area of Expertise:

B.S., Botany, Weber State College, 1979

M.S., Biological Sciences, Northern Arizona University, 1981

Ph.D., Genetics, Iowa State University, 1984

Area of Focus

Plant Breeding and Genetics

Research Interests

Research emphasis is the development and characterization of winter wheat germplasm with improved end-use quality and disease resistance. Supervisory duties include oversight of Lincoln-based USDA research in wheat pathology, sorghum genetics and pathology, and forage grass management, breeding and genetics.

Major Project Activities

  • Wheat molecular and biochemical genetics.
  • Winter wheat germplasm development and regional testing.
  • Grain quality traits.
  • Virus and rust resistance


  • Fakthongphan, J., Graybosch, R. A., & Baenziger, P. S. (2016). Combining Ability for Tolerance to Pre-Harvest Sprouting in Common Wheat (L.). Crop Science, 56(3), 1025-1035.
  • Hystad, S. M., Martin, J. M., Graybosch, R. A., & Giroux, M. J. (2015). Genetic characterization and expression analysis of wheat (Triticum aestivum) line 07OR1074 exhibiting very low polyphenol oxidase (PPO) activity. Theoretical and Applied Genetics, 128(8), 1605-1615.
  • Bartels, M., French, R., Graybosch, R. A., & Tatineni, S. (2016). Triticum mosaic virus exhibits limited population variation yet shows evidence of parallel evolution after replicated serial passage in wheat. Virology, 492, 92-100.
  • Hogg, A. C., Gause, K., Hofer, P., Martin, J. M., Graybosch, R. A., Hansen, L. E., & Giroux, M. J. (2013). Creation of a high-amylose durum wheat through mutagenesis of starch synthase II (SSIIa). Journal of Cereal Science, 57(3), 377-383.
  • Delwiche, S. R., & Graybosch, R. A. (2016). Binary mixtures of waxy wheat and conventional wheat as measured by nir reflectance. Talanta, 146, 496-506.
  • Fakthongphan, J., Bai, G., Amand, P. S., Graybosch, R. A., & Baenziger, P. S. (2015). Identification of markers linked to genes for sprouting tolerance (independent of grain color) in hard white winter wheat (HWWW). Theoretical and Applied Genetics, 129: 419-430.
  • Graybosch, R. A., & Hansen, L. E. (2015). Functionality of chemically modified waxy, partial waxy and wild‐type starches from common wheat.Starch‐Stärke.  doi:10.1002/star.201500241
  • Tatineni, S., McMechan, A. J., Wosula, E. N., Wegulo, S. N., Graybosch, R. A., French, R., & Hein, G. L. (2014). An eriophyid mite-transmitted plant virus contains eight genomic RNA segments with unusual heterogeneity in the nucleocapsid protein. Journal of virology, 88(20), 11834-11845.
  • Rhazi, L., Graybosch, R. A., & Aussenac, T. (2014). Molecular characterization of three transgenic high molecular weight glutenin subunit events in winter wheat. Journal of Cereal Science, 60(3), 631-638.
  • Graybosch, R. A., Amand, P., & Bai, G. (2013). Evaluation of genetic markers for prediction of preharvest sprouting tolerance in hard white winter wheats. Plant Breeding, 132(4), 359-366.
  • Kumssa, T. T., Baenziger, P. S., Rouse, M. N., Guttieri, M., Dweikat, I., Brown-Guedira, G., ... & Poland, J. (2015). Characterization of Stem Rust Resistance in Wheat Cultivar Gage. Crop Science, 55(1), 229-239.
  • Hunt, H. V., Moots, H. M., Graybosch, R. A., Jones, H., Parker, M., Romanova, O., ... & Trafford, K. (2013). Waxy phenotype evolution in the allotetraploid cereal broomcorn millet: mutations at the GBSSI locus in their functional and phylogenetic context. Molecular biology and evolution, 30(1), 109-122.
  • Liu, S., Rudd, J. C., Bai, G., Haley, S. D., Ibrahim, A. M., Xue, Q., ... & St Amand, P. (2014). Molecular markers linked to important genes in hard winter wheat. Crop Science, 54(4), 1304-1321.
  • Tatineni, S., McMechan, A. J., Bartels, M., Hein, G. L., & Graybosch, R. A. (2015). In Vitro Transcripts of Wild-Type and Fluorescent Protein-Tagged Triticum mosaic virus (Family Potyviridae) are Biologically Active in Wheat.Phytopathology, 105(11), 1496-1505.
  • Baenziger, P. S., Graybosch, R. A., Santra, D. K., Regassa, T., Jin, Y., Wegulo, S., ... & Dowell, F. (2014). Registration of ‘Mattern’Waxy (Amylose-free) Winter Wheat. Journal of Plant Registration, 8, 43-48.
  • Tatineni, S., Riethoven, J. J. M., Graybosch, R. A., French, R., & Mitra, A. (2014). Dynamics of small RNA profiles of virus and host origin in wheat cultivars synergistically infected by Wheat streak mosaic virus and Triticum mosaic virus: virus infection caused a drastic shift in the endogenous small RNA profile. PloS one, 9(11), e111577.
  • Graybosch, R. A., Seabourn, B., Chen, Y. R., & Blechl, A. E. (2013). Transgenic Enhancement of High-Molecular-Weight Glutenin Subunit 1Dy10 Concentration: Effects in Wheat Flour Blends and Sponge and Dough Baking 1. Cereal Chemistry, 90(2), 164-168.
  • Byamukama, E., Wegulo, S. N., Tatineni, S., Hein, G. L., Graybosch, R. A., Baenziger, P. S., & French, R. (2014). Quantification of yield loss caused by Triticum mosaic virus and Wheat streak mosaic virus in winter wheat under field conditions. Plant Disease, 98(1), 127-133.
  • Nilthong, S., Graybosch, R. A., & Baenziger, P. S. (2013). Enzyme activity in wheat breeding lines derived from matings of low polyphenol oxidase parents. Euphytica, 190(1), 65-73.
  • Delwiche, S. R., Yang, I. C., & Graybosch, R. A. (2013). Multiple view image analysis of freefalling US wheat grains for damage assessment. Computers and electronics in agriculture, 98, 62-73.
  • Graybosch, R. A., & Peterson, C. J. (2012). Specific adaptation and genetic progress for grain yield in Great Plains hard winter wheats from 1987 to 2010. Crop science, 52(2), 631-643.
  • Nilthong, S., Graybosch, R. A., & Baenziger, P. S. (2012). Inheritance of grain polyphenol oxidase (PPO) activity in multiple wheat (Triticum aestivum L.) genetic backgrounds. Theoretical and Applied Genetics, 125(8), 1705-1715.
  • Graybosch, R. A., & Peterson, C. J. (2010). Genetic improvement in winter wheat yields in the Great Plains of North America, 1959–2008. Crop Science, 50(5), 1882-1890.